Numbered citations not in consecutive order (Nature)

After citation 31 (with no change in page style or breaks) the next citation is listed as 1 (which is missing from the start of the document, citations begin at number 2).

I found all versions of the citation in the document and deleted them, then refreshed and it disappeared from the reference section. When I re-added it it still shows as 1, and next citation is 32.

Style: Nature
Zotero: 5.0.53 latest update installed

Any ideas on how to fix this please?
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