KOReader highlights not importing to EPUB in Zotero

I read EPUBs in KOReader and store the highlights in the sidecar directory. When I import the EPUB into Zotero and open the EPUB in Zotero Reader and attempt to import the highlights into it using File → Import EBook Annotations... I select the correct metadata.epub.lua file for the book, but the import fails:

> “metadata.epub.lua” does not appear to contain any Calibre or KOReader annotations.
>
> If this ebook has been annotated with KOReader, try selecting a “metadata.epub.lua” file directly.

The EPUB was annotated in KOReader and no changes were made to the annotation file, which I can see has all the data as expected. What is causing Zotero Reader to choke? Does it expect a particular folder naming convention for the sidecar directory? It would be ideal to be able to drag-and-drop the metadata.epub.lua file right into the EPUB in Zotero Reader and have the annotations import automatically, is that possible?

Here's my setup:
- KOReader (latest) on a reMarkable Paper Pro Move
- Zotero 8.0.3
- MacOS Tahoe 26.3.

Thanks! (I can provide the EPUB and annotations file, if needed.)
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