Highlights in PDF save selectively

When I open PDFs via Zotero and highlight/annotate, often times I will notice that some, but not all of my highlights save, despite clicking "save" before closing the PDFs. Does anyone have an idea what the reason for this could be?

Some background info: I have a Mac; the PDFs are opened with Preview; my data directory location is set to default; I use data syncing. Let me know if I left out any relevant information.

Thanks in advance!
  • Zotero (before the current Zotero beta, which includes a built-in PDF reader) just launches the PDF in your system PDF reader. After that Zotero isn't involved. So if you save in Preview and then re-open the file immediately and don't see some of the highlights, that wouldn't have anything to do with Zotero.
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